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{ <http://www.tkuhn.ch/bel2nanopub/RARBSnuxzI1C449XwyQbg22Ndc4rN7gysF6E0whBf-9ts#_6> ?p ?o ?g. }
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Early studies established that nuclear translocation of R-Smads is independent of Smad4, whereas translocation of Smad4 after TGF-b signalling seems to require the presence of an activated R-Smad (Hoodless et al., 1999; Liu et al., 1997).
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