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- _2 value "Treatment with the phosphodiesterase (PDE) inhibitors rolipram (a cAMP-specific PDE IV inhibitor), 3-isobutyl-1-methylxanthine (IBMX), erythro-9-(2- hydroxy-3-nonyl)-adenine (EHNA, a PDE II inhibitor), and the b- adrenergic receptor (b-AR) agonist isoproterenol, which increase the intracellular cAMP level, also inhibited the nuclear export of HDAC5 (Fig. 1C andD)." provenance.