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- Study bibliographicCitation "Arnaud, F. (1972) Invert�br�s marins des XII�me et XV�me Exp�ditions Antarctiques Fran�aises en Terre Ad�lie. 9- Pycnogonides. T�thys Suppl., 4: 135-155 (As cited in Gusso, C.C. and Gravina, M.F. (2001) Faunistic and biological traits of some Antarctic pycnogonida. Italian Journal of Zoology,68(4):335-344)" Provenance.